OHNOLOGS

A Repository of Genes Retained from Whole Genome Duplications in the Vertebrate Genomes

Ohnolog families for stickleback (Gasterosteus aculeatus) for criterion: q-score(outgroups) < 0.01 AND q-score(self comparison) < 0.01 from 2R WGD

Family Size	Ohnolog Family
2	mylka	mylk5
3	gad1b, gad1a	zgc:163121	gad2
2	slc6a1b, slc6a1a	slc6a1l
5	APBB1IP(1tomany), apbb1ip	raph1a, raph1b	grb10b, grb10a	GRB14	si:ch211-253b1.3
3	mylpfb, mylpfa	myl2b(1tomany), myl2b(1tomany)	myl7
2	ENSGACG00000001702, tspan7b	tspan7
3	TMEM255B(1tomany)	si:dkey-283j8.1	tmem255a
4	TGFB3(1tomany)	tgfb1a	tgfb3	tgfb2
2	uhrf1bp1l	uhrf1bp1
2	ndel1a, ndel1b	nde1, NDE1(1tomany)
2	e2f7	e2f8
3	abi3b, abi3a	abi1a, abi1b	abi2a, abi2b
3	myom1a, myom1b	si:dkey-78p20.4, si:ch211-270c19.2	myom3
2	spns1	spns3\| spns2
2	slc6a2	SLC6A3
3	APBB2(1tomany), apbb2b	apbb1	APBB3
2	inpp5jb, inpp5ja	inpp5kb, inpp5ka
4	rem2, REM2(1tomany)	gem	rem1	rrad
2	lrrc24, LRRC24(1tomany)	rtn4rl1a, rtn4rl1b
2	nr5a1b, nr5a1a	nr5a2
2	phox2a	phox2bb
2	stxbp1a, stxbp1b	stxbp2
3	zfand5b	zgc:77486	zfand5a
2	gpr137bb(1tomany), GPR137B(1tomany), gpr137bb(1tomany)	gpr137c
4	pax5	pax2b	pax8	pax2a
2	armc1l	armc1
2	rhot1b, rhot1a	rhot2
2	mllt10	MLLT6
3	kcnk2b, kcnk2a	si:dkeyp-115e12.3	kcnk10b
3	si:dkey-242g16.2, zgc:77112	ppp1r3b	ppp1r3ca
3	bag5	bag3	bag4
3	farp1, FARP1(1tomany)	frmd7	farp2
3	zdhhc20a, zdhhc20b	zdhhc2	zdhhc15b
3	sdk2b	si:ch211-148f13.1	sdk1b
2	SPINT1(1tomany), spint1b	spint2
2	ZNF703(1tomany), znf703	ZNF503(1tomany), znf503
2	arl6ip5a, arl6ip5b	praf2
2	atf4a	atf5b
8	fgf11b, fgf11a	fgf13b, fgf13a	fgf5	fgf20b	FGF12(1tomany)	fgf12a	fgf16	fgf9\| fgf14
3	cbfa2t2	CBFA2T3	runx1t1
2	ADAMTS20	adamts9
2	myo9b, si:zfos-588f8.1	myo9ab, MYO9A(1tomany)
3	azin1a, azin1b	ENSGACG00000005287	odc1
2	itm2cb	itm2bb
4	grk5	grk4	grk6	grk5l
3	slc17a6b, slc17a6a	slc17a7b, slc17a7a	slc17a8
3	six9	six1b	six2b
4	si:dkey-202l16.5\| tmc8\| TMC6(1tomany)\| TMC7, tmc6a	tmc2b\| tmc1, tmc2a	tmc4	si:dkey-15b23.3
3	AKT2(1tomany), AKT2(1tomany)	akt3a, akt3b	AKT1
3	DSP(1tomany), dspa	macf1a\| pleca, plecb	si:ch211-165e15.1
2	jag1b, jag1a	jag2b
2	FAM135B	fam135a
6	PLPPR2(1tomany), plppr2a	PLPPR5(1tomany)	plppr5b\| plppr4b	plppr3b	plppr1	plppr3a
5	syt5b, syt5a	syt2a, SYT2(1tomany)	syt8	syt1a	SYT5
2	adarb1a, adarb1b	adarb2(1tomany), adarb2(1tomany)
4	dlx2a	dlx3b	dlx5a	DLX3(1tomany)
7	egfra	erbb3a	erbb3b	erbb4a	erbb4b	EGFR(1tomany)	erbb2
2	lrp1ab, lrp1aa	lrp1bb
9	ENSGACG00000016914, loxl5b	scara5	loxl4	loxa	loxl3b	loxl1	si:ch211-150o23.3	loxl2b	loxl2a
2	cap2	cap1
3	TMEM63B(1tomany), TMEM63B(1tomany)	tmem63a	tmem63c
2	slc27a1a, slc27a1b	slc27a4
2	mmp2	mmp9
2	neto2b, NETO2(1tomany)	neto1, neto1l
2	znf513	znf142
3	slc2a9l1(1tomany), slc2a9l1(1tomany)	slc2a5	slc2a11a
4	ywhabl, SFN	ywhabb	YWHAB(1tomany)	ywhaqb
2	prkar2ab, prkar2aa	si:ch211-272n13.7
7	cpz, CPZ(1tomany)	cpn1	aebp1	CPXM2	cpxm1a	cpe	si:ch1073-459j12.1
2	plod1a	plod2
7	itga11b, itga11a	itga2.2	itga1	itga4	itga10	ENSGACG00000007750, ENSGACG00000007767	itga9
6	marveld2a, marveld2b	si:ch211-13k12.2, ELL3(1tomany)	ell	ELL(1tomany)	ell2	si:dkey-208k22.6
3	SAR1A(1tomany)	sar1ab	sar1b
5	trpm1a(1tomany), trpm1a(1tomany)\| trpm5, trpm1b	si:ch211-173d10.1, trpm4a	TRPM7	trpm2(1tomany), trpm2(1tomany)	TRPM6\| trpm3
2	foxf1	foxf2a
2	s1pr1	lpar1
2	stam	stam2
3	limch1b, limch1a	lmo7a	lmo7b
3	gramd1a	GRAMD1B	si:ch73-93k15.2
4	zfp36l1a	zgc:162730\| zgc:114130	cth1	ENSGACG00000013400
7	slc26a6l, si:ch211-117c9.2	SLC26A9	slc26a6	slc26a5\| slc26a3.2, slc26a3.1	slc26a10	slc26a1	slc26a2
3	reep5	reep6	zgc:101744
4	si:dkeyp-120h9.1	slc7a6	ENSGACG00000016031	slc7a7
2	es1	si:ch211-153b23.5(1tomany), si:ch211-153b23.5(1tomany)
3	C3(1tomany), C3(1tomany), C3(1tomany)	si:dkey-8k3.2	C3(1tomany)
2	stk24b, STK24(1tomany)	stk25b
2	stx1a	stx1b
4	RAB5C(1tomany), rab5c	rab5ab	rab5b	RAB5A(1tomany)
5	kcnq5b, kcnq5a	KCNQ2(1tomany)	kcnq4	kcnq2a	kcnq3
2	gsx2	gsx1
2	dnajc6	gak
2	unkl	unk
2	pik3cg	si:rp71-17i16.5
4	map3k9	map3k10	si:ch211-120p12.3	MAP3K11
3	anp32e(1tomany), anp32e(1tomany)	anp32a	anp32b
2	ktn1	zgc:171356
3	MTSS1(1tomany), MTSS1(1tomany)	MTSS1L(1tomany), mtss1la	mtss1
2	usp44	usp49
5	fhl3b, fhl3a	fhl5	fhl1a	fhl2b	fhl1b
3	srsf1b	srsf9	srsf1a
4	foxo1b, foxo1a	si:ch211-206a7.2, si:dkey-11f5.5	foxo3a, foxo3b	foxo4
5	rab3da, rab3db	rab3c	RAB3A(1tomany)	RAB3A(1tomany)	rab3b
5	WNK2(1tomany), wnk2	wnk1a, wnk1b	WNK4(1tomany)	WNK3	wnk4b
5	hoxc3a	hoxd3a	HOXA3	ENSGACG00000007094	ENSGACG00000003942
3	kdm2bb, kdm2ba	kdm2ab, kdm2aa	zgc:158376
3	nxn	nxnl2	nxnl1(1tomany), nxnl1(1tomany)
3	rxrba, rxrbb	rxraa	rxrgb
2	cacna1ha, cacna1hb	cacna1g
4	pou4f1	zgc:158291	pou4f2	pou4f3
2	si:ch211-278j3.3	rnf19a(1tomany)
3	YTHDF1(1tomany), ythdf1	ythdf3	ythdf2
3	ENSGACG00000012688, ENSGACG00000004618	alox12	alox5a
2	ENSGACG00000018227, atp8b5a	ATP8B2
2	unc45a	unc45b
4	si:dkey-31m14.7, mark4a	MARK1(1tomany), MARK1(1tomany)	mark2b	MARK3
2	zgc:153039(1tomany), zgc:153039(1tomany)	slc12a9
2	ralaa	ralba
2	rtkn2b	rtkn2a
4	grm4, GRM4(1tomany)\| GRM7(1tomany)	grm6a, grm6b	grm8b, grm8a	GRM7(1tomany)
3	nos1apa	nos1apb	si:dkey-34e4.1
4	gabrb3, GABRB3(1tomany)	gabrb2, GABRB2(1tomany)	si:dkey-20i10.4	gabrb4
4	arhgap4b(1tomany), arhgap4b(1tomany), arhgap4a	srgap1a, srgap1b	srgap2	srgap3(1tomany), srgap3(1tomany)
2	plek2	plek
3	cab39l1	cab39l	cab39
2	psmb9a(1tomany), psmb9a(1tomany), psmb9a(1tomany), psmb9a(1tomany), psmb9a(1tomany)\| psmb12(1tomany), psmb12(1tomany), psmb12(1tomany), psmb12(1tomany), psmb12(1tomany)	psmb6
3	zic4\| zic1	zic2a\| zic5	zic6\| zic3
2	dchs1b, dchs1a	si:dkey-30k22.7
2	chchd10	chchd2
2	efhd2	efhd1
2	cav3	cav1
3	vamp3	zgc:92912	vamp2
2	ahcyl1(1tomany), ahcyl1(1tomany)	ahcyl2
2	PEX5L(1tomany), pex5la	pex5
2	fuca2	FUCA1(1tomany), fuca1.2
3	hif1al	epas1b	hif-1a
8	oprd1b(1tomany), oprd1b(1tomany)	OPRK1(1tomany), oprk1	sstr5, SSTR5(1tomany)	SSTR2(1tomany), SSTR2(1tomany)	NPBWR2\| oprl1	sstr3	oprm1	SSTR1
2	mboat2a, mboat2b	mboat4
2	ACTBL2(1tomany), ACTBL2(1tomany)	ACTBL2(1tomany), ACTBL2(1tomany)
4	lmna, LMNA(1tomany)	lmnb2	lmnl3	lmnb1
2	spop	spopla
3	arhgap22, si:dkey-191m6.4	arhgap24	arhgap25
3	asb13b, asb13a.2	asb11\| ASB9	asb5a
4	fgf8a, fgf8b	fgf24	fgf18a	fgf17
3	dpp6a, dpp6b	DPP10(1tomany), DPP10(1tomany), ENSGACG00000001595\| DPP10(1tomany)	ENSGACG00000010877
2	rab11fip4b, rab11fip4a	rab11fip3
3	UNC13B(1tomany), unc13ba	unc13c	UNC13A
2	si:ch211-10a23.2	lgalslb
2	si:dkey-4p15.3	hsp70.3
2	PCBP3(1tomany), pcbp3	pcbp2\| pcbp4, ENSGACG00000006681
5	si:dkey-65j6.2	quo	arhgef40	PLEKHG4B	plekhg4
4	igfbp6a, igfbp6b	igfbp1a\| igfbp3(1tomany), igfbp3(1tomany)\| igfbp1b	igfbp2a, igfbp2b\| igfbp5a	IGFBP4
2	fitm1, FITM1(1tomany)	fitm2
3	fstl1b	fstl4	fstl5
6	xkr7, zgc:198371	xkr6b, XKR6(1tomany)	xkr5	xkr9	xkr8.2\| xkr8.3	zgc:123035
2	rnf150b(1tomany), rnf150a	rnf130
2	sgtb	sgta
3	LRRC4C(1tomany), sc:d0316	lrrc4bb, lrrc4ba	lrrc4.1, lrrc4.2
2	ATXN7L2(1tomany), atxn7l2b	atxn7
3	SLC9A3R2(1tomany)\| SLC9A3R1(1tomany), SLC9A3R2(1tomany), slc9a3r1	pdzk1	pdzd3a
2	osbp2, si:ch211-106a19.1	osbp
2	cyb5r2	cyb5r3
2	casp3a, casp3b(1tomany), casp3b(1tomany), casp3b(1tomany)	casp7
4	ppfia3	ppfia2	ppfia4	PPFIA1
2	smyd1b	smyd2a\| smyd3
2	scinlb, scinla	gsna
2	klhl41a	klhl40b
2	rab44	si:ch211-247i17.1
2	cdc34a, cdc34b	ube2r2
3	gpc5a	ENSGACG00000013942	gpc3
2	si:dkey-36g24.3, adck3	adck4
3	map4k5	map4k3b	map4k2
2	klf18	klf5b
2	zgc:112294	tmem17(1tomany), tmem17(1tomany)
3	onecut3a, ONECUT3(1tomany)	onecutl	onecut1
2	slc25a36b(1tomany), slc25a36b(1tomany)	slc25a33
2	tbc1d22b	tbc1d22a
4	ENSGACG00000004274, ENSGACG00000004284, ENSGACG00000004280, ENSGACG00000004279, ENSGACG00000015361	taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany), taar15(1tomany)	si:dkey-247m21.3	si:ch211-228b4.1
3	si:dkey-112m2.1\| si:dkey-1d7.3, si:dkeyp-14d3.1\| ENSGACG00000012207	TMEM132A	si:dkey-215k6.1
3	sept8a, sept8b	sept10	sept6
4	lhfpl5a, lhfpl5b	LHFPL3(1tomany), lhfpl3	lhfpl4a, LHFPL4(1tomany)	lhfpl4b
2	arhgap21a, arhgap21b	arhgap23a, arhgap23b
3	jph1b, jph1a	jph3, si:ch211-146m13.3	jph2
7	kdrl	PDGFRB\| csf1ra	flt1	kdr\| pdgfra	kitb	csf1rb	flt4
3	PIK3C2G	pik3c2a	pik3c2b(1tomany), pik3c2b(1tomany)
9	kcnj2b, kcnj16\| kcnj2a	KCNJ10(1tomany)\| kcnj9, kcnj10a	KCNJ4(1tomany), KCNJ4(1tomany)	kcnj14	kcnj5\| kcnj1a.4	kcnj11l	kcnj3a	KCNJ15\| KCNJ6	kcnj12
2	kpna4	kpna3
2	NT5C3B(1tomany), NT5C3B(1tomany)	nt5c3a
3	kdm7aa	phf8	phf2
2	gch2	gch1
3	aldh3a2b, aldh3a1\| aldh3a2a	aldh3b2	aldh3b1
4	si:dkey-57c15.1, asic1a	asic4a	ASIC2	asic1c
2	bicc1a	bicc2
2	ctsba	ctsbb
2	SEPT7(1tomany), sept7b	sept15
3	wfikkn2a	si:ch73-359p15.2	wfikkn1
3	chrne	chrng	chrnb1l\| chrnb1
2	angel2	angel1
2	sox10(1tomany), sox10(1tomany)	SOX9(1tomany)\| SOX8(1tomany), sox8a\| SOX9(1tomany)
3	col4a1	col4a3	col4a5
4	sipa1l1, SIPA1L1(1tomany)	sipa1l3	SIPA1	si:dkey-77m16.2
2	si:dkeyp-75h12.2, si:ch211-113e8.3	tesk2
2	si:zfos-632h11.7	col12a1b
2	gpsm2, gpsm2l	gpsm1b
2	ryr1b, ryr1a	RYR3(1tomany), ryr3
2	LDB1(1tomany), ldb1a	ldb2a
2	atf6	ATF6B
2	afg3l2	si:ch1073-174d20.2
2	chpt1	cept1b
8	mmp14a, mmp14b	mmp15a, MMP15(1tomany)	mmp16b, MMP16(1tomany)	mmp11a	mmp20b\| mmp13b	MMP25	mmp24	mmp17a(1tomany), mmp17a(1tomany)
4	scamp2\| scamp5a, scamp2l\| scamp5b	scamp1	scamp4	scamp3
2	zgc:153395	skor1a
3	eomesa	tbr1b	tbx21
2	rela	relb
3	rps6kc1	rps6kl1	snx15
2	ism2b	ism1
3	SS18L1	ss18	ss18l2
3	mogat3a	atp6v1e1a	dgat2\| mogat2
4	chl1b\| nfasca, chl1a	NFASC(1tomany)	nrcama	nadl1.2
2	uck2b, uck2a	uck1
4	phldb2a, phldb2b	phldb1b	phldb1a	PHLDB3
2	yipf2	yipf1
2	ubr1	ubr2
3	si:dkey-261o10.2	dennd1b	si:dkey-76b14.2
2	hmcn2	hmcn1
2	otub1b	OTUB2
3	PXN(1tomany), PXN(1tomany)	si:rp71-15d4.1	TGFB1I1
4	cacna1sb, cacna1sa	cacna1db, cacna1da	cacna1fb, cacna1fa	cacna1c
2	smox	si:dkey-275b16.2
3	ENSGACG00000009272	si:ch73-335m24.2	eva1c
2	prkcdb, prkcda	prkcq
4	pdlim3a	pdlim4	pdlim2	pdlim1
3	hspb15	hspb8\| si:dkey-1k23.3	hspb1
2	tob1a, tob1b	TOB2
3	slc29a1b, slc29a1a	SLC29A2(1tomany)	slc29a2
3	flnca, flncb	flna(1tomany), flna(1tomany)	flnbl
4	ARSI(1tomany), ARSI(1tomany)	ARSB	si:dkey-174i8.1	ARSJ
3	ptbp1b, ptbp1a	ptbp2b, ptbp2a	ptbp3
2	NAV2	nav3
2	CCKBR(1tomany), CCKBR(1tomany)	CCKAR
2	cacng4a, cacng4b	CACNG2\| cacng3b
4	crtc1a	crtc3	CRTC2	crtc1b
3	smad6b	smad7	smad6a
4	sgk1	si:ch211-195b13.1	sgk3	sgk2a
4	HIVEP2(1tomany)	hivep2a	znf831	hivep1
2	ak5	ak1
4	scn1lab, scn1a	scn4aa, scn4ab(1tomany), scn4ab(1tomany)	SCN8A(1tomany), SCN8A(1tomany)	scn12aa
2	si:dkey-153k10.9, si:dkey-5i3.1	si:ch211-163l21.8
2	ndnfl	ndnf
3	prdm1b	prdm1a	prdm1c
2	nkx6.3	nkx6.2
2	pdia3	zgc:100906
2	ehd1b	ehd3
4	celf3b, celf3a	celf4	si:dkey-205h23.2	celf5b
4	GNA11(1tomany), gna11b	GNAQ(1tomany)	gnaq	GNAQ(1tomany)
3	nrbp2b	NRBP1(1tomany)	nrbp1
3	prox1b	prox2	PROX2(1tomany)\| prox1a
3	ap1m1	ap1m3	ap1m2
2	si:dkey-238d18.3	zgc:112160\| ctrb1
3	atl2, ATL2(1tomany)	atl3	atl1
2	trappc6bl	trappc6b
2	axin1, AXIN1(1tomany)	axin2, AXIN2(1tomany)
3	si:zfos-80g12.1	sesn2	sesn1
8	itgb8	itgb2	itgb6	itgb1b	itgb4\| itgb3b	itgb5	itgb3a	itgb1a
2	mief1	mief2
2	narfl	narf
2	rasl11a	rasl11b
4	si:dkey-110k5.6	TBC1D8	tbc1d9	TBC1D9B
2	svila(1tomany), svila(1tomany)	si:dkeyp-94b4.1, si:dkeyp-57f11.2
3	si:dkey-237h12.3, TENM3(1tomany)	tenm4	si:ch211-12m10.1
4	slc25a24, SLC25A24(1tomany)	slc25a23a	slc25a25a	slc25a25b
3	HEY1(1tomany), hey1	heyl	hey2
6	chst7	chst2b	CHST1	zgc:194879	chst3b	si:ch73-62b13.1
2	slc2a6	slc2a8
2	atp8a2	atp8a1
2	megf6b	zgc:158328
2	socs4	socs5
2	ralgapa2	ralgapa1
3	sh3bp1\| arhgap44, si:ch211-114m9.1	ENSGACG00000009145	arhgap17b
2	eps15l1a, EPS15L1(1tomany)	eps15
3	bnip3lb, bnip3la	zgc:73226	bnip4
4	slc25a18	si:dkey-201c1.2	zgc:92113	slc25a22
3	tnk2a	tnk1	tnk2b
4	EPB41(1tomany), epb41a	EPB41L1(1tomany), im:7155308	epb41l3b	si:ch73-18b11.1
2	pdcb, PDC(1tomany)	pdcl
3	thbs4b, thbs4a	si:ch211-43f4.1	thbs3a
2	nid1b(1tomany), nid1a, nid1b(1tomany)	nid2a
2	naca	nacad
3	fcho2	sgip1b	fcho1\| sgip1a
4	smtnl, SMTNL2(1tomany)	SMTN	smtna	smtnb
5	cdh4(1tomany)\| cdh26.1, cdh4(1tomany)	cdh2, CDH2(1tomany)\| dsc2l	cdh13	si:ch211-206i14.2	cdh15
5	grik1b, grik1a	si:ch211-264f5.8, grik5	GRIK3\| GRIK2(1tomany)	grik4	GRIK2(1tomany)
4	cebpg	CEBPE	cebpb	cebpd
2	srsf10b	srsf2a
2	rad9b	rad9a
2	phospho1, PHOSPHO1(1tomany)	phospho2
4	cnksr2a, cnksr2b	cnksr3\| CNKSR3, ipcef1	si:ch211-26b3.4	cnksr1
3	iqgap2	iqgap1	IQGAP3
6	dpysl5a, dpysl5b	dpysl2b, DPYSL2(1tomany)	dpysl3	crmp1	dpys	dpysl4
2	sftpba, sftpbb	psap
3	htra3a, HTRA3(1tomany)	htra1b, htra1a	si:dkey-112g5.12
3	NET1(1tomany), net1	arhgef3l	arhgef3
3	PRSS35(1tomany), prss35	PRSS23	prss23
6	cers2a, cers2b	cers4a, cers4b	cers5	CERS5(1tomany)	cers3b	cers6
3	prkacaa, prkacab	prkacba	prkacbb
3	myhb(1tomany), myhb(1tomany), MYH13(1tomany), MYH13(1tomany), MYH13(1tomany), MYH13(1tomany)	vmhc, smyhc2	myh7bb
2	ccna1	ccna2
2	si:ch211-216l23.1	znf521
2	phkg1b	phkg2
3	epn3a, epn3b	epn1	epn2
3	rfx1b, rfx1a	rfx2	rfx3
2	agpat4	agpat3
3	RBMS1(1tomany), RBMS1(1tomany)	rbms3(1tomany), rbms3(1tomany)	rbms2b, rbms2a
4	si:ch211-230g15.5, phc3	phc1	PHC1(1tomany)	phc2b
7	ptger1b, ptger1a	ptger4a, ptger4b	ptger2a	ptgfr	ptger1c	ptger4c	tbxa2r\| ptger3
2	ENSGACG00000008314, hoxa2b	hoxb2a
2	casq1b, casq1a	casq2, CASQ2(1tomany)
3	rnf24	zgc:175214	si:ch1073-392o20.1
2	gtf3aa, gtf3ab	si:dkey-156n14.3
3	ric8b	si:dkey-94f20.4	ric8a
3	zgc:101559	rab33ba	rab33a
4	rorcb, rorca	roraa	rorc	rorb
2	hhatlb, hhatla	hhat
3	itga6b, itga6a	itga3a	ITGA7
3	tbx3a	tbx2b	ENSGACG00000013575
2	mlxip	si:ch1073-189o9.1
2	srcap	ep400
2	gigyf1	gigfy2
2	wee2	wee1
3	dennd4a, DENND4A(1tomany)	DENND4C	dennd4b
3	ctsd	ren	zgc:63831
8	inaa, inab	nefla\| nefma, neflb	desma(1tomany), desma(1tomany)	si:dkey-222n6.2	viml	si:dkey-27m7.4	ENSGACG00000014542	gfap
2	ano10b	ano10a
2	ARHGAP39	zgc:92107
2	ehd2a, ehd2b	ehd4
2	phip	brwd3
3	si:dkey-178e17.1	slc25a1b	slc25a1a
3	tas1r3	ENSGACG00000006762, ENSGACG00000006510, ENSGACG00000006534, ENSGACG00000006517, ENSGACG00000006529, ENSGACG00000006525, ENSGACG00000006506, ENSGACG00000006536	si:ch1073-155h21.1(1tomany), si:ch1073-155h21.1(1tomany)
4	ctdsplb, ctdspla	ctdsp2	ctdsp1	CTDSP1(1tomany)
2	FMN2(1tomany), fmn2b	fmn1, si:ch1073-209e23.1
2	cdip1	si:ch211-202h22.9
2	wnt5a	wnt5b
4	atp2a2b, atp2a2a	atp2a1l, atp2a1	si:dkey-28b4.8	atp2a3
4	zgc:171558, rapgef4	rapgef5b, rapgef5a	rapgef3	rapgefl1
4	chd7	chd9	chd6	chd8
2	UHRF2	uhrf1
2	fes	fer
2	sos1	sos2
4	lhx8a	lhx8b	lhx6	si:ch211-236k19.2
4	hoxb13a(1tomany), hoxb13a(1tomany)	HOXA11(1tomany), hoxa11a\| hoxa13a\| HOXA10(1tomany)\| hoxa9a	hoxd10a\| hoxd11a\| hoxd12a	hoxc10a\| hoxc12b\| hoxc13b\| hoxc11a
2	plekho1a, plekho1b	plekho2
2	myef2	hnrnpm
3	kif5ab, kif5aa	kif5c, KIF5C(1tomany)	kif5ba, KIF5B(1tomany)
2	PER2(1tomany), per2	per1a
2	CPSF7	cpsf6
2	ehmt1a, ehmt1b	EHMT2
2	sppl2	SPPL2A
2	papss2a	papss1
2	si:ch211-236h17.3, CHST13(1tomany)	chst11
3	osbpl7, OSBPL7(1tomany)	osbpl3b, osbpl3a	osbpl6
2	zgc:153929	ccdc64(1tomany)
4	EBF1(1tomany), EBF1(1tomany)	ebf3a	EBF4	ebf2
2	ogdhb, ogdha	OGDHL
3	rad23ab	rad23aa	rad23b(1tomany)
2	BEST3	best1(1tomany), best1(1tomany)
2	myo1cb, myo1ca	myo1ha, myo1hb
2	DYNC1I2(1tomany), DYNC1I2(1tomany)	DYNC1I1
2	TMCC1(1tomany), tmcc2\| tmcc1b	tmcc3
2	aak1a, aak1b	bmp2k
3	tspan9b, tspan9a	CD53(1tomany)\| zgc:64051	tspan4b
5	ca15b, si:ch211-173d10.4	ca6(1tomany), ca6(1tomany)	ca12	car15	ca14
3	ttc9c	si:ch211-259k16.3	ttc9b
2	nfkb1	nfkb2
3	ENSGACG00000017628, ENSGACG00000011492, ENSGACG00000011467, ENSGACG00000018488	slc5a7a(1tomany), slc5a7a(1tomany)	ENSGACG00000018817
3	pabpc1a, pabpc1b	pabpc4	pabpc1l
3	gnrhr2(1tomany), gnrhr2(1tomany)	gnrhr4	ENSGACG00000003123
3	ca2, cahz\| ca8	ca7	ca5a
2	klf3	klf12b
2	CSGALNACT1(1tomany), CSGALNACT1(1tomany)	csgalnact2
2	dcp1a	dcp1b
2	rims4	rims3
3	si:dkeyp-89c11.1	ENSGACG00000000439	zgc:101553
3	slc4a1b, slc4a1a	slc4a2b, slc4a2a(1tomany), slc4a2a(1tomany)	SLC4A3
10	MID1(1tomany), mid1	trim46b, trim46a	trim55b, trim55a	trim59\| trim54	trim13	trim36	trim9	mid2	trim63b	trim101
2	THRA(1tomany), THRA(1tomany)	thrb
3	ENSGACG00000022246	ENSGACG00000022273	ENSGACG00000022313
2	nfkbiab, nfkbiaa	nfkbie
3	lrtm2b, lrtm2a	lrrc38a, lrrc38b	LRTM1
3	slc19a3a	slc19a2	im:7144090
2	si:dkey-87k14.1, flrt2	flrt1b
4	tfa	mfi2	otomp	zgc:172271
4	ENSGACG00000011331\| ENSGACG00000011309, si:dkey-33m11.8	zgc:171509	ENSGACG00000003239, ENSGACG00000003292, ENSGACG00000003238	try(1tomany), try(1tomany)
2	si:ch1073-284b18.2	LIN28B
3	cnfn	plac8l1	plac8.1
2	th2	th
2	nell2a	NELL1
2	fbxo25	fbxo32
3	reep1	reep2	reep3b
3	arhgef18a, arhgef18b	AKAP13(1tomany)	arhgef1a
4	atp2b1a, atp2b1b	atp2b3b	atp2b2	atp2b4
3	ENSGACG00000001884, ENSGACG00000001881	serpinc1	SERPINB1(1tomany), SERPINB1(1tomany), SERPINB1(1tomany)
2	foxj3, FOXJ3(1tomany)	foxj2
2	zgc:154055	mtmr9
7	ghrhra\| adcyap1r1a\| pth1ra, ghrhrb\| adcyap1r1b	vipr1b, vipr1a	pth2r	pth1rb	ghrhrl	gcgra	SCTR
2	extl2	ext2
2	HAND1	hand2
3	elmo3	elmo1	elmo2
2	CASKIN2(1tomany), CASKIN2(1tomany)	si:dkeyp-9d4.3, si:dkey-34b22.2
2	dopey2	dopey1
3	OPN4(1tomany), opn4a	opn4xb, opn4xa	rrh
2	clint1a	CLINT1(1tomany)
3	ACTC1(1tomany)	ACTC1(1tomany)	si:ch73-187m15.4
3	ppp3ccb, ppp3cca	ppp3ca, ppp3cb	PPP3CA(1tomany)
2	rab11al	rab11bb
2	ahr2(1tomany), ahr2(1tomany)	ahrrb
2	tdh2	tdh
3	TSHZ1(1tomany), tshz1	TSHZ2	tshz3b
2	wnt3	wnt3a
2	rhoac\| RHOC(1tomany), RHOC(1tomany)	rhoab(1tomany), rhoab(1tomany)
3	si:ch211-284f22.3	zgc:114173	vps37c
2	fstb	fstl3
2	bokb	boka
3	gnai3\| gnat2	gnat1	gnai2a
3	met	mst1rb	MST1R(1tomany)
2	cntn3a.2\| cntn3a.1, cntn4\| cntn2\| CNTN3(1tomany)	cntn5
2	irs2a, irs2b	irs4a
2	anks1aa, anks1ab	anks1b
2	si:dkeyp-27e10.3, si:ch211-1e14.1	kiaa1549la, kiaa1549lb
2	setb	seta
6	slc1a3b, slc1a3a	slc1a8a\| slc1a7b, slc1a8b, slc1a7a	slc1a5	slc1a6	slc1a1	slc1a4
2	plekha8	plekha3
2	kpna1	kpna6
3	dnajb12a, dnajb12b	dnajb14	dnajc18
3	SEMA7A	sema7a	si:ch211-113g11.6
2	si:dkeyp-118h3.5	gpr63
2	robo2\| ROBO3(1tomany)\| robo1, ROBO1(1tomany)\| ROBO2(1tomany)	robo3\| robo4
4	hs3st2, HS3ST2(1tomany)	si:ch211-216b21.2\| hs3st3b1b	hs3st3l	HS3ST6
5	b4galt2	b4galt3	b4galt4	si:dkey-199f5.8	b4galt1l
2	grhpra	grhprb
4	sh3bgrl2	sh3bgrl3	sh3bgr	sh3bgrl
2	brms1lb	brms1
3	clcn2b(1tomany)	clcn1b	clcn2a
2	hbp1	ENSGACG00000019192
2	rhbdl3	rhbdl1
3	tp53bp2a, tp53bp2b	ppp1r13bb	ppp1r13ba
2	GRAMD2(1tomany), GRAMD2(1tomany)	GRAMD3(1tomany), GRAMD3(1tomany)
2	DUSP8	dusp16
2	tmed9	tmed4
3	nucb2b	nucb2a	nucb1
3	serinc2, serinc2l	si:ch73-267c23.10	serinc1
3	cyp17a2	cyp17a1	cyp21a2
3	nkain4	nkain2	nkain1
4	hdac4(1tomany), hdac4(1tomany)	hdac5	hdac7b	hdac9b
4	gpc1b, gpc1a	gpc6a, gpc6b	gpc4	gpc2
2	kmt2d	KMT2C
3	usp2b	usp2a	usp21
3	cacna1ba, cacna1bb	cacna1eb, CACNA1E(1tomany)	cacna1aa
3	IHH(1tomany), ihha	dhh	shha
3	kcnk3b, kcnk3a	kcnk9	kcnk15
6	ENSGACG00000009380, slc16a7	slc16a5a(1tomany), slc16a5a(1tomany)	SLC16A3(1tomany), SLC16A3(1tomany)	slc16a1b	SLC16A7	slc16a1a
2	nap1l1	nap1l4a
3	fxr2	fxr1	fmr1
2	frem2a, frem2b	frem3
2	st6galnac5b, st6galnac3\| st6galnac5a	st6galnac6\| st6galnac4
2	gpat4, GPAT4(1tomany)	agpat9l
2	psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany), psmb8a(1tomany)	psmb5
2	CSNK1G2(1tomany), csnk1g2a	csnk1g1
2	xpo1b	xpo1a
2	RARB(1tomany)	RARA
3	mafgb	mafk	maff
4	MAST1(1tomany), MAST1(1tomany)	mast3b, MAST3(1tomany)	MAST4	mast2
5	NFE2L1	nfe2	nfe2l1a	nfe2l3	nfe2l2a
2	prkab1b, prkab1a	PRKAB2
3	tnika(1tomany), tnika(1tomany)	mink1	si:zfos-433a6.1
2	gatad2ab, GATAD2A(1tomany)	gatad2b
2	calr	calr3a
5	rhcg, rhcgb	rh50(1tomany), rh50(1tomany)	rhd	rhbg	rhag
3	ano5b, ano3\| ano5a	ano6	ANO4
2	cyfip2	cyfip1
3	sall3b, sall3a	sall4	sall1a
2	cerk	zgc:158263
3	slc41a2a, slc41a2b	ENSGACG00000003711	SLC41A3\| slc41a1
6	esrrga	ESRRB	esrrgb	ENSGACG00000010561	esrrd	esrra
2	ccnt2a, ccnt2b	ccnt1
4	wasf3b, wasf3a	wasf2	zgc:109889	wasf1
3	arl5a	arl5c	arl8
3	si:ch211-180f4.1	lrrn2	LRRN3
3	LONRF3\| lonrf1	si:ch1073-440b2.1	lonrf1l
2	TKT(1tomany), tkta	tktb
8	ntng2b, NTNG2(1tomany)	si:dkey-220k22.1	ntng1a	NTN5	NTN1(1tomany)	ntn1a	ush2a	ntn2
3	adamts2	ADAMTS14	adamts3
2	ifrd1	ifrd2
3	ap1s3a, ap1s3b	ap1s2(1tomany), ap1s2(1tomany)	ap1s1
2	PPP4R3B	smek1
3	TPD52L2	tpd52l1	tpd52
4	kcnip1b, KCNIP1(1tomany)	kcnip2(1tomany), kcnip2(1tomany)	kcnip3b, KCNIP3(1tomany)	KCNIP4
2	fam167ab	fam167b
3	myo5b	myo5c\| myo5ab	myo5aa
4	pfkfb2a, pfkfb2b	pfkfb4a, pfkfb4b	pfkfb1	pfkfb3
2	myo18aa, myo18ab	MYO18B(1tomany), MYO18B(1tomany)
5	DOCK3(1tomany), dock3	dock1	dock4b	dock5	DOCK2
3	IRF2BP2	irf2bpl	irf2bp1
2	atrx	ENSGACG00000020345
6	GABRG3(1tomany)\| GABRA5(1tomany), GABRG3(1tomany)\| gabra5	GABRA2\| zgc:110204	gabra6a	gabra3	gabra1(1tomany), gabra1(1tomany)\| gabrg2\| gabra6b	si:dkey-155h10.3
3	capn3a	capn3b	capn9
4	gpm6ab	gpm6aa	gpm6bb	gpm6ba
5	clic5b, clic6\| clic5a	clic4	clic3	clic2	clic1
2	THBS1(1tomany), THBS1(1tomany)	thbs2b
4	camk1ga, camk1gb	camk1da, camk1db	camk1a	camk1b
2	ENSGACG00000014978	rbmx
2	bcor	bcorl1
2	valopb	opn1sw2\| opn1lw1
5	ckbb, ckba	ckmt2b(1tomany), ckmt2b(1tomany)	ckmb, ckma	ckmt1	ENSGACG00000019583
2	mgrn1b	si:dkey-49c17.3
2	asap1b	asap2a
3	araf	raf1b	braf
6	cacnb4.1, cacnb4a	cacnb3a	cacnb1	cacnb2b	cacnb3b	cacnb2a
4	egr1	egr3	egr2a	egr2b
3	btbd11b	abtb2b	btbd11a
4	srpk3, SRPK3(1tomany)	srpk1b, srpk1a	si:ch211-220i18.4	srpk2
2	TWF1	twf2b
2	lrrk2	lrrk1
3	TRIP10(1tomany), trip10a	fnbp1b	fnbp1l
2	si:ch211-87m7.2, ENSGACG00000018277	hiat1a
2	ppip5k1a(1tomany), ppip5k1a(1tomany)	ppip5k1b
2	mfn1b, MFN1(1tomany)	mfn2
5	zgc:101583	nipa2\| nipa1	nipal3	NIPAL2	nipal4
2	TNPO2(1tomany), TNPO2(1tomany)	tnpo1
3	slc22a15	slc22a2(1tomany), slc22a2(1tomany)	slc22a16
2	MCTP1(1tomany), mctp1b	mctp2a, mctp2b
2	slc25a12(1tomany), slc25a12(1tomany)	slc25a13
3	clcn5b	clcn4	clcn3
3	atp6v0a1a, atp6v0a1b	atp6v0a2b, atp6v0a2a	tcirg1b
3	txlnbb, txlnba	txlna	txlng
2	cacna2d4b, CACNA2D4(1tomany)	CACNA2D3(1tomany), cacna2d3
2	ENG	tgfbr3
8	ugt5a1, ENSGACG00000016578, ugt5b4, ENSGACG00000016569	ugt5e1, ENSGACG00000019954, ENSGACG00000019953	ugt1b2	si:ch73-334d15.1	ugt5d1	ENSGACG00000013879, ENSGACG00000013898, ENSGACG00000013890, ENSGACG00000013874	ugt1a1	ugt8
4	tbc1d10aa	tbc1d10ab	tbc1d10c	tbc1d10b
4	slc12a7a, slc12a7b	slc12a5a, slc12a5b	si:cabz01090193.1	slc12a4
2	fam117ab, fam117aa	fam117ba
2	scarb2	wu:fi18e02
2	nsfl1c	ubxn2a
4	c1ql4b, c1ql4a	c1ql3b	c1ql3b	c1ql2
3	stard14	STARD10	si:dkeyp-110e4.11
2	samd4a	si:ch1073-111c8.3
3	ENSGACG00000009596, ENSGACG00000009583	itih5\| itih2	ITIH4(1tomany), ITIH4(1tomany)
3	sema6d, sema6e	sema6bb, sema6ba	SEMA6A(1tomany), SEMA6A(1tomany)
3	arhgef25b, ARHGEF25(1tomany)	kalrnb, kalrna	trioa, triob
3	ampd3a, ampd3b	ampd2b	ampd1
2	sltm	safb
5	zfr2	zfr	ilf3b	strbp	ilf3a
2	ccdc146	cfap58
2	foxn4	foxn1
5	CDH18\| cdh10a\| cdh12a	CDH22	CDH19\| CDH7	cdh11\| cdh5	cdh8
2	ATP6V0D2	atp6v0d1
2	sec23ip	ddhd2
2	exoc6b	exoc6
3	sept9a, sept9b	sept3	sept12
2	rbl2	rbl1
2	klhl2	klhl3
2	ZMIZ1(1tomany), zmiz1b	ZMIZ2
3	twist3(1tomany)	twist3b	twist2
4	rlbp1a, rlbp1b	ttpa	ttpal	clvs2
2	frs2b	frs3
2	zgc:103586	cda
2	epb41l4b	epb41l5
5	atp1b2b, atp1b2a	atp1b1b, atp1b1a	ATP4B	atp1b3a	atp1b4
5	adgra1a, ADGRA1(1tomany)	lgi2a\| ADGRA3, lgi2b	lgi1b, lgi1a	lgi3	adgra2
2	si:ch211-282j22.3	edem3
3	ABHD17A(1tomany), ABHD17A(1tomany)	abhd17b	abhd17c
3	mat2ab, mat2aa	mat1a, MAT1A(1tomany)	mat2al
2	adamts6	adamts10
2	mapk6	mapk4
3	atp1a3a, atp1a3b	atp1a2a	atp1a1a.4
7	cyp24a1	cyp27c1	cyp11a2	cyp27a1.2	cyp27b1	si:dkey-91i10.3	cyp11c1
2	lmtk2	aatkb
2	dmxl2	DMXL1
2	rab27b(1tomany), rab27b(1tomany)	rab27a
2	cobll1b	cobl
2	dennd5b(1tomany), dennd5b(1tomany)	dennd5a
3	kdm4ab, kdm4aa	kdm4c	kdm4b
3	notch1b, notch1a	notch2	notch3
3	zgc:56235, vdac3	vdac2	vdac1
4	si:dkey-22i16.2, cnga1	cnga5	cnga4	cnga3a
2	adamts15b\| adamts15b, adamts15a	ADAMTS1\| ADAMTS5
2	fblim1	zyx
3	ctbp2a, CTBP2(1tomany)	ctbp1	zgc:136929
2	serp2	zgc:85858
2	slc39a6	slc39a10
2	RASSF10(1tomany), rassf10a	rassf9
2	SPHK1(1tomany), SPHK1(1tomany)	sphk2
2	thsd7ba	thsd7ab
4	fosaa, si:ch211-153j24.3\| fosab	fosl2	fosb	fosl1a
2	dyrk2	zgc:172180
3	elovl1a	elovl1b	elovl7a
2	tiam1a	tiam2a
4	rgmd	rgmb	rgma	hfe2
2	cntnap5a	cntnap1
2	anxa4	anxa1a
2	ap2b1	ap1b1
3	fam20cb(1tomany), fam20cb(1tomany)	fam20a	ENSGACG00000008362
3	TCF12(1tomany), tcf12	tcf3a, tcf3b	TCF4
4	tor2a	tor1	tor4ab	tor3a
2	tmub1	tmub2
2	fem1c	fem1a
3	ENSGACG00000011228	si:ch211-22d5.2	nim1k
4	actn2b	actn3a	actn1	ACTN4
2	prmt8b, PRMT8(1tomany)	prmt1
2	LMBRD2(1tomany)	lmbrd2b
3	cpne1	cpne3(1tomany), cpne3(1tomany), cpne3(1tomany), cpne3(1tomany)	cpne2
2	rbpms2b	RBPMS
12	EPHB3(1tomany), ephb3a	ephb2a, ephb2b	ephb4a, ephb4b	ephb1, EPHB1(1tomany)	epha3	epha4b	epha6	EPHA5	ephb6	EPHA2	epha4a	epha7
3	bin1a, bin1b	bin2b	amph
2	u2surp	zgc:163098
4	dvl3a	dixdc1a	dvl2	dvl1a
2	mfge8b, mfge8a	edil3
4	txnbb, tnxba	tnr\| tnw	tnc	TNC(1tomany)
5	mitfb, mitfa	TFE3(1tomany)	tfec	tfe3b	tfeb
3	dgkab, dgkaa	dgkg	dgkb
2	nuak1b	nuak2
3	si:ch211-173b16.3, ank1b	ank3b, ank3a	ANK2(1tomany), ank2b
3	mkrn4	mkrn2	mkrn1
2	ctsa	ENSGACG00000015897
3	mcoln3a, MCOLN3(1tomany)	mcoln1a, mcoln1b	MCOLN2
2	wdfy1	wdfy2
2	tmem55a	tmem55bb
2	hip1rb	hip1
2	lysmd3	lysmd4
3	mep1a.2	si:ch73-40a17.3	mep1b
6	FBLN1(1tomany), fbln1	fbln2, FBLN2(1tomany)	efemp2a	EFEMP1	fbln5	FBLN7
3	sorbs2a	sorbs1	sorbs3
2	auts2a	fbrsl1
3	arfip2b	arfip2a	arfip1
2	ccnjl	ccnj
4	dusp4	dusp5	dusp2	dusp1
4	tfap2e	tfap2d\| tfap2b	tfap2a	tfap2c
5	si:ch73-368j24.3\| hist1h4l\| ENSGACG00000013913	si:ch1073-153i20.1	zgc:163040	si:dkey-23a13.15	HIST2H4A\| HIST1H4C\| si:ch73-368j24.4(1tomany)\| zgc:163040\| si:ch73-368j24.4(1tomany)\| si:ch1073-159d7.1\| si:dkey-108k21.18\| si:ch73-368j24.2
3	ndst1a	ndst2a	ndst3
2	fam102aa, fam102ab	fam102ba, fam102bb
2	adcy2a, adcy2b(1tomany), adcy2b(1tomany)	adcy7
2	im:6904482, si:ch211-288d18.1	usp6nl
4	atcaya, atcayb	bnip2	BNIPL	PRUNE2
4	lrp8, LRP8(1tomany)	ldlra, ldlrb	vldlr	lrp1ba
2	si:ch73-141f14.1	agtpbp1
3	ZSWIM6(1tomany), ZSWIM6(1tomany)	zswim5	zswim8
2	atp10d	atp10a
3	gipc1	gipc2	gipc3
2	sostdc1b	sost
2	GAL3ST4	gal3st1a
2	cox6a2	cox6a1
2	stox2b, stox2a	stox1
4	rph3ab, RPH3A(1tomany)	DOC2A(1tomany), si:dkey-93n13.3	doc2b	doc2d
2	rgs6(1tomany), rgs6(1tomany)	rgs7a
2	si:ch211-69m14.1	frmd3
4	gdf9	GDF10	bmp3	bmp15
2	ncf2	noxa1
3	celsr2	celsr3	celsr1a
9	arvcfa, arvcfb	CTNND1(1tomany), ctnnd1	CTNND2(1tomany), ctnnd2b	pkp4, PKP4(1tomany)	pkp1b	pkp3a	PKP3(1tomany)	pkp1a	pkp2
2	ypel3	ypel2a
2	egfl6	npnt
2	ENSGACG00000012709, ENSGACG00000016754	grap2a
2	RNF213	rnf213b
2	cdr2l, CDR2L(1tomany)	cdr2a, si:ch1073-272o11.3
3	LPCAT1(1tomany), lpcat1	lpcat4	lpcat2
4	coro1cb	coro6	coro1ca	coro1a
4	KCNC3(1tomany), kcnc3a	kcnc4	kcnc2	kcnc1b
3	itga5, ITGA5(1tomany)	itgav	itga8
2	pex11b	pex11a
3	tacc1, TACC1(1tomany)	tacc3	TACC2
3	LPIN2(1tomany), lipin2	si:ch73-21k16.1, lpin1	zgc:123305
2	camkk1a, camkk1b	si:ch73-62l21.1
2	CRHR2	crhr1
3	ENSGACG00000022247	ENSGACG00000022192	ENSGACG00000022169
3	CCNG2(1tomany), ccni\| ccng2	ccni2	ccng1
2	zgc:175280(1tomany)\| slc7a1, slc7a2\| ENSGACG00000018362, SLC7A1(1tomany)	slc7a3a
2	rprd1b	rprd1a
3	HOMER1(1tomany), homer1b	homer3, HOMER3(1tomany)	HOMER2
6	lmod2a, lmod2b	tmod1	tmod2	lmod1b	lmod3	tmod4
2	CHUK(1tomany), chuk